Hi,
Which Exchange org/system is the target of the MX record for Company2.com? I am assuming the SMTP address space of your Exchange org is org2.company2.com, is this correct? Is this also the primary address in the Default recipient policy?
The following KB article will be of use to you: http://support.microsoft.com/?kbid=321721
Basically the solution calls for you to configure an additional recipient policy.
1. Your default recipient policy will have the primary SMTP address space of org2.company2.com and the checkbox "This Exchange Org is responsible for all mail delivered to this address" will be checked
2. The additional Recipient policy will have company2.com as the primary SMTP entry but this time the check box "This Exchange Org is responsible for all mail delivered to this address" is NOT checked.
You will also need to change the order of the recipient policies so that the default has less priority than the newly created one, you will also need to create/apply a suitable LDAP filter for the new recipient policy so that it is applied appropriately
The procedure above will ensure your email server does not think that it is responsible for all email bound for the company2.com SMTP name space and will forward any messages out for which a mailbox/address cannot be found in your Exchange org.
If you have any further questions don't hesitate to reply to my post.
Regards
Ben Hoffman
Microsoft MVP - Windows Server System Exchange